I think (based on your wife's foot) the high speeds and wind resistance is killing the mpg. I have a 2024 Limited non hybrid with air dam removed, OEM Tonneau cover installed and consistently get 29-31 mpg.
Driving conditions are: 41 miles each way with a 4-5 degree steady elevation change over...
My Limited lives in the NC Outer Banks and all that I carry with me is an air compressor to air back up, a long rope, front and back recovery points, soft shackles and traction boards.
My 2019 4Runner TRD Off Road is an overlanding beast and when driving on the beach I have never had to deploy...